Stephen H. Schneider, a Stanford University biologist on the vanguard of climate-change research for four decades, who argued eloquently on human culpability in global warming and willingly threw himself into the political fray to explain and defend the scientific evidence, has died. He was 65.

Schneider had a heart attack Monday while flying to London from a science meeting in Stockholm, according to Stanford spokesman Dan Stober.

"Steve Schneider helped the world understand that the burning of fossils had altered the chemistry of Earth's atmosphere, and that this change … had led to a discernible human influence on our planet's climate," said Benjamin D. Santer, a leading climate researcher at Lawrence Livermore National Laboratory, who described his colleague as the Carl Sagan of climate science.

Santer and Schneider were among the scientists who served on the international panel that shared the 2007 Nobel Peace Prize with former Vice President Al Gore, who in a statement Monday called Schneider a "prolific researcher and author … and a wonderful communicator" whose contributions to the advancement of climate science will be "sorely missed."

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